Life sciences · Preprint
arXiv · September 21, 2026
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The quadratic scaling of Transformer self-attention has driven the adoption of sub-quadratic Selective State Space Models (SSMs) like Mamba, which compress past context into a fixed-size recurrent hidden state. This strict informational bottleneck raises a foundational question for mechanistic interpretability: do SSMs and Transformers learn fundamentally distinct latent representations? In this work, we employ Sparse Autoencoders (SAEs) to conduct a large-scale, feature-level correspondence analysis between Mamba-130m and Pythia-70m over a 10-million token corpus. Contrary to hypotheses predicting widespread architectural divergence, we find no evidence of systematic representational divergence between architectures: across the observed Jaccard distribution, 99.98% of Mamba features cluster toward the upper alignment boundary, providing preliminary feature-level support for the Universality Hypothesis. We further identify and qualitatively characterize this microscopic fraction (0.02%) of diverging features, finding patterns consistent with the hypothesis that the recurrent bottleneck selectively limits the parsing of rigid syntax rather than broad semantic ontology. We demonstrate that while Pythia's unconstrained attention permits the monosemantic decomposition of distinct formatting edge-cases, Mamba is forced to compress unrelated syntactical anomalies into polysemantic "junk drawer" neurons to preserve state capacity. Collectively, these results suggest that architectural routing mechanisms may have negligible impact on core semantic understanding, with representational divergence confined to extreme structural margins.