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Life sciences · Preprint
arXiv · September 16, 2026
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We analyze the learned input-output behavior of GLU-based neurons in large language models (LLMs). We propose a simple analysis method: For each neuron, we compute the cosine similarities between its input (reading) and output (writing) weight vectors. In this scheme, a strong negative cosine similarity indicates the neuron weakens the direction it detects in the residual stream, so we call this a weakening neuron. This allows us to gain a number of novel insights. First, we show that nine different LLMs have similar patterns: weakening neurons appear mostly in late layers whereas their counterparts, (conditional) strengthening neurons, are frequent in early-middle layers. Second, we find that weakening neurons display surprising behavior: even though there are few, they activate often and have a large influence on model behavior. Third, weakening neurons have a strong effect on model output when gate values are negative -- which is surprising since negative gate values are not expected to encode functionality.